Open Access
Issue
BIO Web Conf.
Volume 215, 2026
The International Congress on Natural Resources and Sustainable Development (RENA 2025)
Article Number 01007
Number of page(s) 10
Section Biodiversity and Functioning of Natural Ecosystems
DOI https://doi.org/10.1051/bioconf/202621501007
Published online 04 February 2026
  • J. Vasconez-Gonzalez, H. Alexander-León, M.L. de Noboa-Lasso, J.S. Izquierdo-Condoy, E. Puente-Villamarín, and E. Ortiz-Prado, Scorpionism: a neglected tropical disease with global public health implications, Front Public Health 13, 1603857 (2025). [Google Scholar]
  • J.-A. Vaucel et al., French Scorpionism (Mainland and Oversea Territories): Narrative Review of Scorpion Species, Scorpion Venom, and Envenoming Management, Toxins 14, 719 (2022). [Google Scholar]
  • F. Abroug, L. Ouanes-Besbes, N. Tilouche, and S. Elatrous, Scorpion envenomation: state of the art, Intensive Care Med 46, 401 (2020). [Google Scholar]
  • P. Sousa, M. A. Arnedo, and D. J. Harris, Updated catalogue and taxonomic notes on the Old-World scorpion genus Buthus Leach, 1815 (Scorpiones, Buthidae), Zookeys 15 (2017). [Google Scholar]
  • M.M. Kamel, O. Tanane, R. Saile, and A. Kettani, Fauna and Zoogeography of Scorpions (Arachnida: Scorpions) in Morocco, Ecology, Environment and Conservation 28, S31 (2022). [Google Scholar]
  • M. Abdelmonaim, M. A. el Hidan, O. Touloun, and A. Boumezzough, Spatial relationship between environmental factors and scorpion distribution in Morocco, Journal of Entomology and Zoology Studies 674, 674 (2017). [Google Scholar]
  • F. Salhi, J. P. Dunbar, C. Lawton, J. Hermas, J. A. Oualid, and M. M. Dugon, Geographic distribution of the scorpion fauna in the central Moroccan region of Souss-Massa with potential implications for public health, African Zoology 59, 39 (2024). [Google Scholar]
  • A. Elmourid, S. Boussaa, M. A. El Hidan, O. Amahmid, and O. Touloun, Epidemiological, toxicological and physiopathological characteristics of scorpion stings and their management in Morocco: A literature review, Acta Tropica 239, 106812 (2023). [Google Scholar]
  • capm-sante.ma/uploads/documents/62.pdf?utm_source=chatgpt.com, (n.d.). [Google Scholar]
  • Z. Ruiming, M. Yibao, H. Yawen, D. Zhiyong, W. Yingliang, C. Zhijian, and L. Wenxin, Comparative venom gland transcriptome analysis of the scorpion Lychas mucronatus reveals intraspecific toxic gene diversity and new venomous components, BMC Genomics 11, 452 (2010). [Google Scholar]
  • K. Ammouch, N. Mesmoudi, N. Hammani, J. Galan, A. Moustaghfir, R. Stöcklin, and N. Oukkache, Tackling the burden of envenomation in Africa: advances, challenges, and strategic priorities for enhanced diagnosis and treatment, Front. Trop. Dis. 6, (2025). [Google Scholar]
  • B. Ajdi, A. El Asbahani, M. A. El Hidan, M. Bocquet, L. Falconnet, M. Ait Hamza, A. Elmourid, O. Touloun, H. Boubaker, and P. Bulet, Molecular diversity assessed by MALDI mass spectrometry of two scorpion species venom from two different locations in Morocco, Toxicon 238, 107562 (2024). [Google Scholar]
  • P.R.S. Bencheikh, Scorpion : un drame évitable, (n.d.). [Google Scholar]
  • Boulaouane (Rural Commune, Morocco) - Population Statistics, Charts, Map and Location, https://www.citypopulation.de/en/morocco/grandcasablancasettat/admin/el_jadida/1810901__boulaouane/. [Google Scholar]
  • Quand Partir à Boulaouane ? Climat, Météo et Meilleure Période | Maroc, https://planificateur.a-contresens.net/afrique/maroc/casablanca-settat/boulaouane/2554409.html. [Google Scholar]
  • R. Klesser, M. Husemann, T. Schmitt, P. Sousa, A. Moussi, and J. C. Habel, Molecular biogeography of the Mediterranean Buthus species complex (Scorpiones: Buthidae) at its southern Palaearctic margin, Biol J Linn Soc 133, 166 (2021). [Google Scholar]
  • M. Nazari and R. Hassan, Study on Distribution of Scorpions to Provide Prevention and Interventions in Combating Scorpionism in Poldokhtar County, Lorestan Province, Iran, J Clin Diagn Res 10, LC05 (2016). [Google Scholar]
  • R. Dehghani, A. M. Tehrani, F. Ghadami, and H. Sanaei-Zadeh, Methods for Collecting and Capturing Scorpions: A Review, 4, (2016). [Google Scholar]
  • M. Vachon and R. Stockmann, Contribution À L’étude Des Scorpions Africains Appartenant Au Genre Buthotus Vachon 1949 Et Étude De La Variabilité: RICERCHE SULLA FAUNA DELLA SOMALIA PROMOSSE DALL’ISTITUTO DI ZOOLOGIA E DAL MUSEO ZOOLOGICO DELL’UNIVERSITÀ DI FIRENZE: XXXIII, Monitore Zoologico Italiano. Supplemento 2, 81 (1968). [Google Scholar]
  • W. R. Lourenço, Scorpion incidents, misidentification cases and possible implications for the final interpretation of results, J. Venom. Anim. Toxins Incl. Trop. Dis 22, 21 (2016). [Google Scholar]
  • S. L. Leary, editor , AVMA Guidelines for the Euthanasia of Animals: 2020 Edition, 2020 edition (American Veterinary Medical Association, Schaumburg, IL, 2020). [Google Scholar]
  • N. H. Shomer, K. H. Allen-Worthington, D.L. Hickman, M. Jonnalagadda, J.T. Newsome, A.R. Slate, H. Valentine, A.M. Williams, and M. Wilkinson, Review of Rodent Euthanasia Methods, J Am Assoc Lab Anim Sci 59, 242 (2020). [Google Scholar]
  • D. Marquina, M. Buczek, F. Ronquist, and P. Łukasik, The effect of ethanol concentration on the morphological and molecular preservation of insects for biodiversity studies, PeerJ 9, e10799 (2021). [Google Scholar]
  • S. McGraw, Taxonomy Sample Collection and Preservation., https://www.ecoanalysts.com/sample-collection-and-preservation. [Google Scholar]
  • Linnaeus, The Scorpion Files - Scorpio Maurus (Scorpionidae), https://www.ntnu.no/ub/scorpion-files/s_maurus.php. [Google Scholar]
  • E. Froufe, P. Sousa, P. C. Alves, and D. J. Harris, Genetic diversity within Scorpio maurus (Scorpiones: Scorpionidae) from morocco: Preliminary evidence based on CO1 mitochondrial DNA sequences, Biologia 63, 1157 (2008). [Google Scholar]
  • Ythier, (PDF) A New Species of Buthus Leach, 1815 from the Atlantic Coast of Morocco (Scorpiones: Buthidae), https://www.researchgate.net/publication/374842484_A_new_species_of_Buthus_Leach_1815_from_the_Atlantic_coast_of_Morocco_Scorpiones_Buthidae. [Google Scholar]
  • D. El Ghoubali, S. Pirro, S. Sehli, M. Merzouki, N. A. Idrissi, L. Belyamani, S. Hamdi, and H. Ghazal, The complete genome sequence of Androctonus mauritanicus, the Moroccan black thick-tailed scorpion, Biodivers Genomes 2022, 10.56179/001c.55548 (2022). [Google Scholar]
  • R. El Oufir, I. Semlali, M. Idrissi, A. Soulaymani, S. Benlarabi, A. Khattabi, M. Ait Moh, and R. Soulaymani Bencheikh, Scorpion sting: a public health problem in El Kelaa des Sraghna (Morocco), J. Venom. Anim. Toxins Incl. Trop. Dis 14, 258 (2008). [Google Scholar]
  • I. Hilal, S. Khourcha, A. Safi, A. Hmyene, S. Asnawi, I. Othman, R. Stöcklin, and N. Oukkache, Comparative Proteomic Analysis of the Venoms from the Most Dangerous Scorpions in Morocco: Androctonus mauritanicus and Buthus occitanus, Life 13, 1133 (2023). [Google Scholar]
  • J. Heurtault, M. Goyffon, and R. Stockmann, La fonction venimeuse et les venins, Annales de l’Institut Pasteur / Actualités 10, 147 (1999). [Google Scholar]
  • M. A. Omran, The Scorpion and its venom (Review article), Serket 13, 51 (2012). [Google Scholar]
  • J.-P. Chippaux, ENVENIMATIONS ET INTOXICATIONS PAR LES ANIMAUX VENIMEUX OU VÉNÉNEUX II. ENVENIMATIONS PAR VIPERIDAE, (n.d.). [Google Scholar]
  • W. R. Lourenço, Why does the number of dangerous species of scorpions increase? The particular case of the genus Leiurus Ehrenberg (Buthidae) in Africa, J. Venom. Anim. Toxins Incl. Trop. Dis 26, e20200041 (2020). [Google Scholar]
  • J.-P. Chippaux and M. Goyffon, ENVENIMATIONS ET INTOXICATIONS PAR LES ANIMAUX VENIMEUX OU VÉNÉNEUX I. GÉNÉRALITÉS, (n.d.). [Google Scholar]

Current usage metrics show cumulative count of Article Views (full-text article views including HTML views, PDF and ePub downloads, according to the available data) and Abstracts Views on Vision4Press platform.

Data correspond to usage on the plateform after 2015. The current usage metrics is available 48-96 hours after online publication and is updated daily on week days.

Initial download of the metrics may take a while.